Harlequins have strengthened their front-row resources for next season with the signing of England prop Paul Doran-Jones on a two-year contract.
The 27-year-old joins from Northampton Saints where he has been playing for the past two seasons.
He played for England at U18 as well as Ireland at U19 and U21 levels before finally declaring for England.
He made his international debut against Argentina during the 2009 November Tests and has since made three further appearances including being part of the team that won the Six Nations in 2011.
He was a member of the England squad that toured South Africa in June 2012.
"While we were disappointed to lose James Johnston we recognise the need to have an experienced player to come in to supplement the existing tighthead talent we have at the club in Will Collier and Kyle Sinckler. With that in mind we are very fortunate to be able to bring in a player of Paul's ability," said Harlequins director of rugby, Conor O'Shea.
"Not only is Paul talented, at his age he will only get better. He comes from a fabulous club and has been coached by one of the best forwards coaches in the country so I have no doubt he will add greatly to what we have at Harlequins.
"We look forward to welcoming him to The Stoop at the start of next season."